Origins dives into the thrilling battles of the inter-war and early-war period. This expansion matches biplanes against fast monoplanes in combat over Spain, China, and Poland. Nationalist CR.32s face off against "Red" I-15s over Madrid. Soviet volunteers fly "Chaikas" against Japanese "Claudes" above China. Bf 109s dive into streams of British Wellingtons, Halifaxes, and Manchesters while defending battleships in Brest.

New Aircraft range from pre-war German warplanes (such as the He 46 and He 51) to Polish fighters and bombers (such as the PZL P.11) and American Curtiss fighters (such as the CW-21 Interceptor). In addition to new aircraft and scenarios, Origins features a whole campaign system, recreating two days of the German and Italian maximum effort against Malta in 1942. Titled Malta Spitfires, the campaign simulates the tough attrition battles over the islands.